Including fossils in phylogeny: a glimpse into the evolution of the superfamily Evanioidea (Hymenoptera: Apocrita) under tip-dating and the fossilized birth–death process

Abstract Using a fossilized birth–death model, new phylogeny of the superfamily Evanioidea (including ensign wasps, nightshade wasps and hatchet wasps) is proposed, with estimates divergence times for its constitutive families corroborating monophyly Evanioidea. Additionally, our Bayesian analyses demonstrate †Anomopterellidae, †Othniodellithidae, †Andreneliidae, Aulacidae, Gasteruptiida Evaniidae, whereas †Praeaulacidae †Baissidae appear to be paraphyletic. Vectevania vetula Hyptiogastrites electrinus are transferred Aulacidae. We estimate time in Late Triassic (~203 Mya). three othniodellithid described figured from mid-Cretaceous Burmese amber as genus Keratodellitha, species: Keratodellitha anubis sp. nov., basilisci nov. kirin also document temporal shift relative species richness between Ichneumonoidea